Driving home late at night during a heavy rainstorm, Glen experiences car trouble. Near where his car gets stuck, he spots a house, knocks on the door and is greeted by an oddly friendly middle-aged man, Arthur, and his younger wife, Cyndi. The strange couple pours him a drink, and then more drinks, followed by an unexpected offer that Glen can’t refuse. Elsewhere, a young woman, Katie, is feeling emotionally weighed down by a secret romantic arrangement that feels like a textbook case of gaslighting. And at the same time, in a nondescript research facility, medical professional Julie begins questioning her role in a bizarre experiment, fearing that she’s doing more harm than good.

 故事發生在2008年北京奧運會前夕，參加過朝鮮戰爭的退伍老兵張大川（楊新鳴 飾），與兒子一家生活在中國南方的一個小鎮上，雖然同住一個屋檐下但關系并不親密。一天，大川接到老戰友去世的電話，于是他決定獨自前往北京，參加戰友的葬禮。這個計劃，被正值青春叛逆期的孫子小松（胡昌霖 飾）發現，小松懇求和威脅爺爺帶他一起去。爭執之后，從未單獨出過遠門的祖孫倆上路了.......

　　一個神秘爬行動物的到來，把溫馨的動物城攪動得天翻地覆。面對全新的城市危機，警官兔朱迪（金妮弗·古德溫 Ginnifer Goodwin 配音）與狐尼克（杰森·貝特曼 Jason Bateman 配音）將繼續攜手為保衛動物城而奔波。在追捕行動中，這對老搭檔不僅要揭開新角色的神秘面紗，還要前往被迷霧籠罩的新領域，探索未知的地下黑市，一場瘋狂動物城的全新冒險即將展開……