In the year 2961, the time is after humanity and nature has recovered the land. A hunter named Cygnus is called to rise above his duty. He provides for Last Arc, a once nomadic band of survivors in need of food and water that is now growing scarce. The answer must be found before a group of outlandish Heretics descend upon them. Cygnus must voyage across the treacherous landscape to defend his people. Sent by Nova, the matriarch of the band, she acts based on her vision for Cygnus to find a seed of hope. The future of Last Arc is for him to discover, Cygnus ventures into a hostile landscape in search of an answer for his people. On this journey of encountering many traps and dangers, Cygnus discovers what has been hunting him is his identity.

　　故事發生在某個毗鄰原始森林的小鎮，一家伐木公司正在林中作業，這時白雪飄然落下，作業暫時停止，然而隨即便發生了殘酷血腥的事件。與此同時，警長保羅·希爾德（凱文·杜蘭德 Kevin Durand 飾）和新來的搭檔唐尼剛剛敷衍完幾個報警事件，隨后驅車來到前妻蘇珊（Bianca Kajlich 飾）的家中，接走兒子亞當（Ethan Khusidman 飾）回自家居住。一夜無事，次日一早，保羅在門口發現碩大的腳印，而且腳印遍布小鎮主干道，一直延伸到森林深處。巨大的腳印在小鎮鬧得人心惶惶，保羅起初絕不相信妖精怪物一類的傳說，但是恐怖血腥的事件接二連三發生，讓小鎮陷入前所未有的恐慌之中。　　危機漸次逼近……

拳擊手大卓因意外受傷告別了擂臺，在小區當了一個保安。在照顧患有癡呆癥父親的過程中，大卓漸漸理解了父親多年來對他的愛，父子間童年時的心結被解開。為彌補父親曾經的遺憾，大卓重拾信心，再一次站到了擂臺之上。

　　故事背景設置在不遠的未來世界，在414區的人型機器人殖民地中，私家偵探大衛·卡邁克爾(皮爾斯飾)受雇于癡迷商業世界的財閥馬龍·維特(費米爾飾)去尋找后者叛逆的女兒。卡邁克爾需要與高度智能的機器人(魯茨飾)通力合作，而在這個過程中，關于414區和“機器人之城”起源的罪行也浮出了水面。