在美國陸軍游騎兵選拔的最后階段，一支精英團隊的訓練演習變成了與一種難以想象的威脅之間的生存之戰。

　　After breaking out of a rehabilitation zoo, Frankie, a squirrel with all the angles, returns to his forest. He learns from his dim-witted best friend, Cody, a porcupine and street-con, that while he was in lock-up, the entire forest was picked clean by The A.C.O.R.N.S. Institute (Alternative Cosmetic Oils for Regenerative Natural Skin-care). No acorn was left behind. With winter just around the corner, it's a potentially devastating blow to all forest critters. So, Frankie has no choice but to bust into the brand new, state-of-the-art, rodent-proof A.C.O.R.N.S. Processing Plant and take back all of the acorns! Though a seemingly impossible heist, success would not only save the lives of his forest friends, but it would also give Frankie a chance to win back the love of his life, Lola, a local "show squirrel." Sadly, Lola has been burnt by too many of Frankie's false promises in the past. With the support of Frankie's old mentor, Mr. Bellwood, a wealthy badger, Frankie and Cody assemble a team of top notch "crackers" from all over the forest, including: a bat named Fly Boy, a top notch munitions expert; Raitch, a computer hacking albino snake, who never allows her lack of appendages to get in the way; Edsy a tiny "family frog," whose ability to freeze himself solid for days on end without harm is invaluable; a germophobic French rat named Liam, who can collapse his skeleton to get into any space; Chinese father-daughter pangolins named Pango and Lin, the best diggers in the forest; and Bellwood's hedgehog butler, Oscar who knows how to get his hands on anything the crew needs. With winter knocking at their door, Frankie and his crew have only three days to prepare and execute this unprecedented heist. When they learn that the processing of their forest's acorns has been pushed up a day, Lola's refusal to participate leaves Frankie and his crew one critter short. With everything on the line, Frankie and his crew of Animal Crackerz set out to pull off the biggest heist in animal history!

　　身形健碩的德里克·湯普森（巨石·強森 Dwayne Johnson 飾）是一個作風粗暴的職業冰球手，由于經常打掉別人的牙齒，因此他在場上得了“牙仙”的綽號。正所謂風云變幻，時代更迭，隨著才俊新秀的涌現，德里克也漸漸成為一名過氣的板凳選手。狂放自負的他必定心情極度失落，故而經常用冰冷的語言打碎別人的夢想。　　德里克的女友卡莉（艾什莉·賈德 Ashley Judd 飾）有一雙可愛的兒女，小女兒剛好處在換牙期，對神秘的牙仙還充滿遐想。此時德里克的老毛病又犯了，一心希望向女孩講述牙仙并不存在的真相。他的粗暴做法惹來了麻煩，牙仙大陸的負責人莉莉（朱麗·安德魯斯 Julie Andrews 飾）將他變為一個身長翅膀、手持魔杖、衣著短裙的牙仙女，在接下來的兩周里，他將被迫以這身裝扮為孩子們帶去祝福……

　　哈蒙德（理查德?阿滕伯勒 Richard Attenborough  飾）立志要建立一個非同尋常的公園：恐龍將是這個公園的主角。他把眾多科學家收歸旗下，利用琥珀里面困住的遠古蚊子體內的血液，提取出恐龍的基因信息，利用這些信息培育繁殖恐龍。結果如愿以償，他把怒布拉島建立成了一個恐龍公園，堅信可以從中賺取大錢。然而，科學家們則憂心忡忡。　　不幸的事情果然發生了。雖然公園有電腦系統管理，但卻因為被員工破壞而造成了無法挽救的失控：所有的恐龍逃出了控制區，人們紛紛逃竄卻逃不過恐龍的魔爪。恐龍自相殘殺，人們亦死難無數，最后幸存者寥寥，只得四人逃出生天。怒布拉島上空彌漫著恐怖的氣息。