困于七年前一樁意外的兩人再度相遇，美云（辛芷蕾 飾）與葆樹（張頌文 飾）開始了一段愛恨交織、扭曲而痛苦的共生關系。在美云試圖借其峰（馮紹峰 飾）展望新的未來時，卻發現自己仍深陷舊日漩渦。那些關于情與債的執念，再度將所有人推向命運的十字路口……

　　冷戰時，全球兩大特工組織——俄羅斯“中心”與英國“圓場”明爭暗斗。特工遇害，高層震怒，責成“圓場”挖出“地鼠”。懷疑對象共有5人，即圓場上任“老總”命名的鍋匠，裁縫，士兵，窮人，乞丐。其中，金盆洗手的“乞丐”喬治·史邁利(加里·奧德曼 Gary Oldman 飾)本來有重大嫌疑。某日，行動組-剝頭皮組頭目的彼得·吉勒姆(本尼迪克特·康伯巴奇 Benedict Cumberbatch 飾)與英國內閣辦公室諜報總監拉康排除了史邁利的嫌疑，令其重新出山，負責挖出“地鼠”。于是，史邁利開始分別走訪故人，調查“圓場”老大阿勒萊恩，裁縫-比爾·海頓(科林·費斯 Colin Firth 飾)，士兵-掌管東歐事務的資深間諜羅埃·布蘭德，窮人-后勤部門點路燈組組長托比·伊斯特哈斯。一時間，圓場疑云密布，人人自危，究竟誰才是“地鼠”……　　本片本片根據約翰·勒卡雷的同名小說改編。片名來自英國民間的“鵝媽媽童謠集”。

　　The boisterous, arrogant professor Challenger, a reputed biologist and anthropologist, dares the London Zoological Society to mount an expedition to verify his spectacular claim, without physical proof, that his previous expedition to the Amazonian basin found live dinosaurs. Apart from him and his 'socialite' counterpart, professor Summerlee, it consists of experienced discoverer Lord Roxbury, the young reporter Ed Malone -who got publicly struck down with Challenger's umbrella at his arrival- and Jennifer Holmes, Malone's news agency's boss's daughter, essentially as conditions for putting up the money. In Brazil they are joined by Jennifer's brother David and local 'guide' Manuel Gomez. They soon discover the dinosaurs are real and dangerous, like giant spiders, but loose their helicopter and thus are desperate for a way down from the isolated plateau. They learn Roxbury knew about the fate of Burton White, an explorer whose diary they find, thus presume to be dead, in search of diamonds, and confirmation of the local tribe being the lethal guardians of the plateau's secrets, but also get surprising help...

　　A young mad genius attempts to 'hack the human mind' in order to fix humanity.